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Figshare: figures and data from published and unpublished studies 
iNaturalist: an incomplete, but growing, list of collection records
Bionomia: digitized specimens deposited in museum collections
ORCID, Google Scholar and Research Gate: Links to publications, publication statistics 
Publons: Review and editorial contributions
Presentations can be found here
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Genbank accessions can be found here. 

Please click on the images below for a link to the publication.  Preprints of all papers will be made available when the manuscript has been accepted for publication.

Forthcoming

Standardized terminology and visual atlas of the external morphology and terminalia for the genus Scaptomyza (Diptera: Drosophilidae)

Distribution and Taxonomy of Endemic and Introduced Drosophilidae in Hawaii

Insect Radiations on Islands: Biogeographic Pattern and Evolutionary Process in Hawaiian Insects
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Active Learning Strategies for Biodiversity Science.  Bioscience

Left.  A Molecular Phylogeny of the Endemic Hawaiian Scatella
(Diptera: Ephydridae)

2020

Right.  Evolution of reproductive behavior. Genetics 214: 49-73.  [PDF]

Genetic variability among native and introduced strains of the parasitic nematode Deladenus siricidicola  Journal of Invertebrate Pathology [PDF]

Evidence for both sequential mutations and recombination in the evolution of kdr alleles in Aedes aegypti. PLoS Neglected Tropical Diseases [PDF]

2019

Below.  A standardized nomenclature and atlas of the male terminalia of Drosophila melanogaster. right  [preprint]

Right, top. Niche conservatism predominates in adaptive radiation: comparing the diversification of Hawaiian arthropods using ecological niche modelling. [PDF]
Right, bottom. Cytochrome oxidase I sequences from northern and southern California suggest cryptic Baetis (Ephemeroptera: Baetidae) species. Western North American Naturalist 79(2): 204-218. [PDF]

2018

​Above: Days of Future Past: Hawaiian Drosophila as an Evolutionary Model Clade.  Bioessays [PDF]

​Left: Phylogeny of the genus Drosophila.  Flybook [PDF]
Access the recommendation on F1000
The Genus Leucophenga (Diptera: Drosophilidae), part VIII: Twenty-one Species from the Oriental Region, with Morphological and Molecular Evidence. Zootaxa [PDF]

2017

Above: Temporal patterns of genetic diversity in Baetis tricaudatus (Ephemeroptera: Baetidae) from the Russian River, Northern California.  Freshwater Science [PDF]

Right: Peterson, MG, O'Grady, PM, and Resh, VH.  Phylogeographic comparison of five large-bodied aquatic insect species across the
western United States.  Freshwater Science [PDF]
STUDENT DISSERTATION (co-advised with Vince Resh): Stauffer, NJ. 2017. Spatial and Temporal Variability in Benthic Invertebrate Assemblages and Population Genetics in a Lake and Stream System. University of California, Berkeley. 

2016

Above.  Studies in Hawaiian Diptera III: New Distributional Records for Endemic Canacidae and a New Species of Procanace.  ​Biodiversity Data Journal  [PDF]
​Left.  Multiple colonizations of the Hawaiian Islands by the family Dolichopodidae (Diptera), with a phylogeny of the Eurynogaster complex.  PeerJ [PDF | preprint]
​

2015

Above.  Community assembly on isolated islands: Macroecology meets evolution.   Global Ecology and Biogeography [PDF]
Right.  The effects of introduced species, habitat characteristics, and land use on Hawaiian anchialine pool biota.  Hydrobiologia [PDF]
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​Drosophilidae (Diptera).  State of Hawaii, Division of Forestry and Wildlife, Wildlife Programs Fact Sheets. [Figshare]
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Procanace (Diptera, Canacidae).  State of Hawaii, Division of Forestry and Wildlife, Wildlife Programs Fact Sheets. [Figshare]
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Dicranomyia (Diptera, Limoniidae).  State of Hawaii, Division of Forestry and Wildlife, Wildlife Programs Fact Sheets. [Figshare]
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Scatella (Diptera, Ephyldridae).  State of Hawaii, Division of Forestry and Wildlife, Wildlife Programs Fact Sheets. [Figshare]

2014

Phylogenetics of the antopocerus-modified tarsus clade of Hawaiian Drosophila: Diversification across the Hawaiian Islands.
PLoS ONE [
PDF]
Microbial interactions and the ecology and evolution of Hawaiian Drosophilidae. Frontiers in Microbiology [PDF]
Diversification in Hawaiian long-legged flies (Diptera: Dolichopodidae: Campsicnemus): Biogeographic Isolation and Ecological Adaptation.  Molecular Phylogenetics and Evolution [PDF]
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Evolutionary Insights into cospeciation and host switching in Neotropical batflies (Diptera: Streblidae). [Figshare]  
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Studies in Hawaiian DIptera I: New distributional records for endemic Asteia (Asteiidae).  Biodiversity Data Journal 2:e1010.  [PDF]
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O'Grady Material Deposited in the Ambrose Monell Cryo Collection, 1998-2002. 
[Figshare]
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Studies in Hawaiian Diptera II: New distributional records for endemic Scatella (Ephydridae).  Biodiversity Data Journal 2:e1110. [PDF]
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Catalog of Drosophila species deposited in the Ambrose Monell Cryo Collection by W. B. Heed. [Figshare]  
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Predicting impacts of sea level rise for cultural and natural resources in five national park units on on the Big Island of Hawaii.  Technical Report 133 CESU (University of California at Berkeley and National Park Service. 40 pp. [PDF]
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STUDENT DISSERTATION: Marrack, EC.  2014.  Modeling the Effects of Sea Level Rise and Introduced Species on Hawaiian Anchialine Pool Ecosystems. University of California, Berkeley. 

2013

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Diversification and Dispersal of the Hawaiian Drosophilidae: the evolution of Scaptomyza. Molecular Phylogenetics & Evolution [PDF].  
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O'Grady Lab Collections. [Figshare] 
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Molecular phylogeny and biogeography of the Hawaiian craneflies Dicranomyia (Diptera: Limoniidae). PLoS One [PDF]
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Historical biogeography and ecological opportunity in the adaptive radiation of native Hawaiian leafhoppers (Cicadellidae: Nesophrosyne). Journal of Biogeography [PDF]
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Drosophila suzukii invasion of Hawaii and Oregon assessed by divergence population genetics. [Figshare] 

2012

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Fungal diversity associated with Hawaiian Drosophila host plants. PLoS One [PDF]
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Diversity and phylogenetic relationships of Wolbachia in Drosophila and other native Hawaiian insects.  Fly [PDF]
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Host plants shape insect diversity: origins, diversity, and host plant associations of the native Hawaiian leafhoppers (Cicadellidae: Nesophrosyne).  Molecular Phylogenetics & Evolution [PDF]
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Left.  Monophyly, divergence times, and evolution of host plant use inferred from a revised phylogeny of the Drosophila repleta species group. Molecular Phylogenetics & Evolution [PDF]
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Book Review: Untangling Phylogenetic Networks.  Taxon [PDF]
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Book Review: Retrograde biogeography. Taxon [PDF]
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Estimating divergence dates and substitution rates in the Drosophila phylogeny. Molecular Biology & Evolution [PDF]
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STUDENT DISSERTATION.  Bennett, GM.  2012.  Origins, Diversity, and Diversification of the Native Hawaiian Leafhoppers (Hemiptera: Cicadellidae: Nesophrosyne) And Their Obligate Endosymbionts.  University of California, Berkeley.  174 pages.  [PDF]
Right.  Rapid morphological, behavioral, and ecological evolution in Drosophila: Comparisons between the endemic Hawaiian Drosophila and the cactophilic repleta species group. In: Rapidly Evolving Genes and Genetic Systems  (RS Singh, J Xu and RJ Kulanthial, eds). Oxford University Press. [PDF]
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2011

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Phylogenetic and ecological relationships of the Hawaiian Drosophila inferred by mitochondrial DNA analysis.  Molecular Phylogenetics & Evolution [PDF]
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Phylogenetic relationships in the spoon tarsus subgroup of Hawaiian Drosophila; Conflict and concordance between gene trees. Molecular Phylogenetics & Evolution [PDF]
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Review of the native Hawaiian leafhopper genus Nesophrosyne (Hemiptera: Cicadellidae: Deltocephalinae) with descriptions of eight new species associated with Broussaisia arguta.  Zootaxa [PDF]
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STUDENT DISSERTATION.  Lapoint, RT.  2011.  Diversification in the Hawaiian Drosophila. University of California, Berkeley.  148 pages.   [PDF]
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2010

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A plea for character-based DNA barcoding: Insights from the Drosophilidae. Molecular Phylogenetics & Evolution [PDF]
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​Microbial community profiles enhance detection of groundwater contamination from leaking waste disposal sites.  Water Resources Research [PDF]
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Morphological variation in the forelegs of Hawaiian Drosophilidae.  I The AMC clade.  Journal of Morphology  [PDF]
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Taxonomic relationships within the endemic Hawaiian Drosophilidae.  Records of the Hawaii Biological Survey [PDF]
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Whither Drosophila?  Genetics [PDF]
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Morphological and molecular evidence to support the synonymy of Emperoptera Grimshaw with Campsicnemus Halliday (Diptera: Dolichopodidae). Records of the Hawaii Biological Survey [PDF]


2009

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​Left: Spread of an introduced vector-borne banana virus in Hawai‘i. Molecular Ecology [PDF]
Phylogenetic taxonomy in Drosophila: Problems and prospects.  Fly [PDF] 
Revision of the modified mouthparts species group of Hawaiian Drosophila (Diptera: Drosophilidae): The ceratostoma, freycinetiae, semifuscata, and setiger subgroups and unplaced species. University of California Publications in Entomology [PDF]
Review of the spoon tarsus subgroup of Hawaiian Drosophila  (Drosophilidae: Diptera), with a description of one new species. Zootaxa [PDF]
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Drosophila. In: Encyclopedia of Islands (Gillespie, RG and Clague, D, eds.) [PDF]
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Drosophila melanogaster.  In: Encyclopedia of Insects, volume 2 (Resh, V & Cardé, R, eds.) [PDF]
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New immigrant Drosophilidae in Hawaii, and a checklist of the established immigrant species.  Proceedings of the Hawaiian Entomological Society [PDF]

2008

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Polytene chromosome maps of 11 Drosophila species: The order of genomic scaffolds inferred from genetic and physical maps. Genetics [PDF] ​
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Out of Hawaii: The biogeographic history of the genus Scaptomyza (Diptera: Drosophilidae). Biology Letters [PDF]

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STUDENT THESIS.  Manendo, T.  2008.  A Phylogenetic Analysis of North American Lasius ants based on mitochondrial and nuclear DNA. University of Vermont. 59 pages.  [PDF]
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Evolution of genes and genomes on the Drosophila phylogeny. Nature  [PDF]​​

Using whole genome presence/absence data to untangle function in 12 Drosophila genomes.  Fly [PDF]

Species delimitation in the Drosophila aldrichi subcluster (Diptera: Drosophilidae) using DNA sequences.  Zootaxa  [PDF]

Comment on the proposed conservation of usage of Drosophila Fallén, 1823 (Insecta, Diptera).  Bulletin of Zoological Nomenclature  [PDF]

The potential and peril of the supertree approach: A response to van der Linde and Houle.  Insect Systematics and Evolution [PDF]

Reproductive ecology of Drosophila. Functional Ecology [PDF]

Mitochondrial phylogeny of the endemic Hawaiian craneflies (Diptera, Limoniidae, Dicranomyia): Implications for biogeography and species formation.  Molecular Phylogenetics & Evolution  [PDF]

A review of the endemic Hawaiian Drosophilidae and their host plants. Zootaxa  [PDF]

New combinations in Hawaiian Drosophila and Scaptomyza (Diptera: Drosophilidae). Zootaxa  [PDF]

Revision of the nudidrosophila and ateledrosophila species groups of Hawaiian Drosophila (Diptera: Drosophilidae) with descriptions of twenty-two new species. Systematic Entomology  [PDF]
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Drosophila: A Guide to Species Identification and Use.  Academic Press, London.
2007                                                
Drosophila biology in the genomic age.  Genetics  [PDF] 

​2006                                                
Leafhopper Keys: Hawaiian Subfamily Key and Endemic Leafhoppers, HNVP, Kipuka Puaulu. Taken from Tree of Life Web Project, 28th August, 2007. [Tree of Life] 

Using microbial community profiles for delineation and long-term monitoring of a landfill leachate-contaminated plume.  Geological Society of America, Philadelphia, Pennsylvania.

A subgroup structure for the modified mouthparts group of Hawaiian Drosophila.  Proceedings of the Hawaiian Entomological Society [PDF]

Unreported chironomid diversity in Hawaii and implications for ecological monitoring.  Pacific Branch of the Entomological Society of America, Kihei, Maui, Hawaii. 

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Evolutionary genetics of reproductive behavior in Drosophila: connecting the dots.  Annual Review of Genetics [PDF]
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2005                                                
Systematics and geographical distribution of the Drosophila longicornis species complex (Diptera: Drosophilidae: repleta species group).  Zootaxa [PDF]

Book review of: A Primer of Genome Science, second edition. Cladistics [PDF]

Phylogeny and age of diversification of the planitibia species group of Hawaiian Drosophila.  Molecular Phylogenetics & Evolution   [PDF]

2004                                                
Phylogenetic relationships within the haleakalae species group inferred by molecular and morphological characters (Diptera: Drosophilidae).  Bishop Museum Bulletins in Entomology  [PDF]

2003                                                
Hierarchical structure in the Drosophila mojavensis cluster (Diptera: Drosophilidae).  Hereditas [PDF]

Revision of the Drosophila mimica subgroup, with descriptions of ten new species.  Annals of the Entomological Society of America [PDF]

The placement of the Engiscaptomyza, Grimshawomyia, and Titanochaeta, three clades of endemic Hawaiian Drosophilidae.   Zootaxa [PDF]

Drosophila melanogaster.  In: Encyclopedia of Insects (Resh, V & Cardé, R, eds.)  Academic Press

Proposal for Drosophila as a model system for comparative genomics.  White Paper submitted to NIH 10 June 2003.

The evolution of the Hom-C homeobox cluster in the Dipteran family Drosophilidae. Insect Molecular Biology [PDF]
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2002

23 [B]  Zilversmit, M, O’Grady, PM, and DeSalle, R.  2002.  Shallow Genomics, Phylogenetics, and Evolution in the Family Drosophilidae.  In Pacific Symposium on Biocomputing (Altman, R, Dunker, AK, Hunter, L, Lauderdale, K, Klein, T, eds). pages 512-523.  [PDF]

22 [A]  Remsen, J, and O'Grady, PM.  2002.  Phylogeny of Drosophilidae (Diptera), with comments on combined analysis and character support.  Molecular Phylogenetics and Evolution 24(2): 248-263. [PDF]

21 [A]  O’Grady, PM, Vela, D, and Rafael, V.  2002.  Zygothrica desallei: A new species of Drosophilidae (Diptera) from Ecuador. Annals of the Entomological Society of America 31(3): 314-315. [PDF | Figures]

20 [A]  O’Grady, PM, Beardsley, JW, and Perreira, WD.  2002.  New records for introduced Drosophilidae (Diptera) in Hawai'i. Bishop Museum Occasional Papers 65: 34-35.  [PDF]
19 [A]  O'Grady, PM, Durando, CM, Heed, WB, Wasserman, M, Etges, W, and DeSalle, R.  2002.  Genetic divergence within the Drosophila mayaguana subcluster, a closely related triad of Caribbean species in the repleta species group.  Hereditas: 136(3): 240-226. [PDF | Data, Tables and Figures]

18 [A]  O’Grady, PM, and Kidwell, MG.  2002.  Phylogeny of the subgenus Sophophora (Diptera: Drosophilidae) inferred using the 28S, alcohol dehydrogenase, and cytochrome oxidase II genes.  Molecular Phylogenetics and Evolution: 22(3): 442-453.  [PDF | Data, Tables and Figures]

17 [A]  O'Grady, PM.  2002.  Notes on the nomenclature of the endemic Hawaiian Drosophilidae.  Bishop Museum Occasional Papers 65: 36-40. [PDF | Figures]

16 [B]  O’Grady, PM, Remsen, J, and Gatesy, JE.  2002.  Partitioning of multiple data sets in phylogenetic analysis.  In: Methods and Tools in Biosciences and Medicine:  Techniques in Molecular Evolution and Systematics (DeSalle, R, Giribet, G & Wheeler, W, eds.).  Birkhauser Verlag, Berlin.

15 [B]  O’Grady, PM.  2002.  Species to genera: Phylogenetic inference in the Hawaiian Drosophilidae.  Pp 17-30.  In: Molecular Systematics and Evolution: Theory and Practice (DeSalle, R, Giribet, G. & Wheeler, W., eds.).  Birkhauser Verlag, Berlin.

2001

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14 [C]  Zilversmit, M.  O'Grady, PM,  Russello, M, and DeSalle, R.  2001.  High Throughput Sequencing Protocols for a Survey of Genomic Characters in the Family Drosophilidae.  Drosophila Information Service 84:199-201.  [PDF]

13 [A]  O’Grady, PM,Val, FC, Hardy, DE, and Kaneshiro, KY.   2001.  The rustica species group of Hawaiian Drosophila (Diptera: Drosophilidae).  Pan-Pacific Entomologist 77(4): 254-260. [PDF | Figures]

12 [A]  O’Grady, PM, Baker, RH, Durando, CM, Etges, WJ, DeSalle, R.  2001.  Polytene chromosomes as indicators of phylogeny in several species groups of Drosophila.  BMC Evolutionary Biology 1:6.  [PDF | Journal]

11 [F]  O’Grady, PM.  2001.  A phylogenetic catalog of the Hawaiian Drosophilidae clade (Diptera: Drosophilidae). [site no longer maintained]

10 [A]  Hardy, DE, Kaneshiro, KY, Val, FC, and O’Grady, PM.  2001.  Review of the haleakalae species group of Hawaiian Drosophila (Diptera: Drosophilidae).  Bishop Museum Bulletin in Entomology 9: 1-88.  Bishop Museum Press.

9 [A]  Etges, WJ, Armella, MA, O'Grady, PM, and Heed, WB.  2001.  Two new species of Drosophila (Diptera; Drosophilidae) in the repleta group from Mexico.  Annals of the Entomological Society of America 94(1): 16-20. [PDF | Figures]

8 [C]  Bonacum, J, DeSalle, R, O’Grady, PM, Olivera, D, Wintermute, J, and Zilversmit, M.  2001.  New nuclear and mitochondrial primers for systematics and comparative genomics in Drosophilidae.  Drosophila Information Service 84:201-204. [PDF]

2000

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7 [B]  O’Grady, PM, and DeSalle, R.  2000.  How the fruit fly changed (some of) its spots.  Current Biology 10(2): R75-77. [PDF | Figures]

6 [A]  Heed WB, and O’Grady, PM.  2000.  Drosophila maya, a new Neotropical member of the Drosophila obscura species group.  Journal of the New York Entomological Society  108(1-2): 97-104.  [PDF | Figures]

1999

5 [A]  O’Grady, PM.  1999.  Reevaluation of phylogeny in the Drosophila  obscura species group.  Molecular Phylogenetics and Evolution 12(2): 124-139. [PDF | Data, Tables and Figures]

4 [A]  Gatesy, JE, O’Grady, PM, and Baker, RH.  1999.  Corroboration among data sets in simultaneous analysis: hidden support for phylogenetic relationships among higher level Artiodactyl taxa.  Cladistics 15: 271-313. [PDF]

1998

3 [F]  O'Grady, PM.  1998.  Phylogenetic relationships of flies in family Drosophilidae inferred by combined analysis of morphological and molecular characters.  Ph.D. Dissertation, University of Arizona. [PDF]

2 [A]  O’Grady, PM, Clark, JB, and Kidwell, MG.  1998.  Phylogeny of the Drosophila saltans species group based on combined analysis of nuclear and mitochondrial DNA sequences.  Molecular Biology and Evolution 15(6): 656-664. [PDF | Data, Tables and Figures]

1997

1 [A]  Carareto, CMA, Kim, W, Wojciechowski, MF, O'Grady, PM, Prochorova, AV, Silva, JC, and Kidwell, MG.  1997.  Testing transposable elements as genetic drive mechanisms using Drosophila P element constructs as a model system.  Genetica 101: 13-34. [PDF]
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